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Pearson Edexcel 4MA1 IGCSE  ·  Topic G6.13  ·  Higher tier

G6.13 Capture–Recapture

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01 — Explanation

What's the proportion the whole method relies on?

The assumption that the fraction of marked individuals found in a second sample mirrors the fraction marked in the whole population:

RC=MN\frac{R}{C} = \frac{M}{N}

where MM = marked in the first sample, CC = size of the second sample, RR = marked ones recaptured in it, and NN = the (unknown) total population you're estimating.

Exam Tip: set up this proportion every time rather than memorising a rearranged formula — "part of sample CC over whole of sample CC equals part of population NN over whole of population NN" avoids ever inverting the wrong pair.

02 — Explanation

How do I estimate the population?

Substitute the three known values and cross-multiply to solve for NN.

90 birds ringed (M=90M=90); second sample of 120 (C=120C=120) includes 20 ringed (R=20R=20):

20120=90N  ⟹  20N=90×120  ⟹  N=1080020=540\frac{20}{120}=\frac{90}{N} \implies 20N = 90\times120 \implies N = \frac{10800}{20}=540

03 — Explanation

What assumptions do I need to state, and why do they matter?

Four specific ideas are examinable, not vague ones like "it's just an estimate":

  • No mark loss — tags/dye didn't fall off or fade.
  • Random mixing — the marked individuals spread back out evenly, so the second sample is representative.
  • Closed population — no births, deaths, immigration or emigration between samples.
  • No change to survival/behaviour — the mark doesn't make the animal easier or harder to recatch.

Common Mistake: a vague answer like "the sample was random" scores nothing — say specifically what must stay true (e.g. "the tags did not fall off").

If marks are lost, RR comes out artificially low, which inflates NN (since N=M×CRN=\frac{M\times C}{R} — a smaller RR in the denominator makes NN bigger). Being able to explain this direction of error, not just state the assumption, is what separates a Grade 7 answer from a Grade 6 one.

04 — Explanation

How do I work backwards to find M or R?

The same proportion rearranges to solve for any of the four variables — just substitute what's unknown as XX and solve.

C=50C=50, R=32R=32, N=125N=125, find MM:

3250=M125  ⟹  M=32×12550=80\frac{32}{50}=\frac{M}{125} \implies M = \frac{32\times125}{50}=80

05 — Explanation

How do I handle an algebraic version of the problem?

Grade 9 variants replace numbers with expressions, usually linked by a second piece of information (like "TT is one-fifth of NN"). Substitute everything into the same proportion and solve as normal.

A scientist tags TT lions; the next day finds T−10T-10 lions, 4 of which are tagged; TT is 15\frac15 of the total NN (so N=5TN=5T). Find NN.

  1. RC=MN⇒4T−10=T5T=15\dfrac{R}{C}=\dfrac{M}{N} \Rightarrow \dfrac{4}{T-10}=\dfrac{T}{5T}=\dfrac15.
  2. Cross-multiply: 4×5=T−10⇒20=T−10⇒T=304\times5 = T-10 \Rightarrow 20=T-10 \Rightarrow T=30.
  3. N=5T=150N=5T=150.

06 — Worked example

Worked exam-style question

A park ranger marks 400 fish and releases them. The next day she catches 60 fish, of which 3 are marked. Estimate the total fish population, and state one assumption your estimate relies on.

  1. 360=400N⇒3N=400×60=24000⇒N=8000\dfrac{3}{60}=\dfrac{400}{N} \Rightarrow 3N = 400\times60=24000 \Rightarrow N=8000.
  2. Assumption: the marked fish mixed back thoroughly and randomly with the rest of the population before the second sample was taken (any one specific, named assumption from the list above is acceptable).

Population estimates must always round to a sensible whole number — a population of "9333.3 jelly beans" should be rounded, since individuals can't be fractional.